[1] Univ Queensland, Sch Biol Sci, Australian Res Council Ctr Excellence Environm De, Brisbane, Qld 4072, Australia
[2] Indonesian Inst Sci, Purwodadi Bot Garden, Jalan Surabaya Malang Km 65, Pasuruan 67163, East Java, Indonesia
[3] Borneo Futures Project, Country Woods 306,Jalan WR Supratman, Ciputat 15412, Tangerang, Indonesia
[4] Ctr Int Forestry Res, POB 0113 BOCBD, Bogor 16000, Indonesia
[5] Univ Kent, Sch Anthropol & Conservat, DICE, Canterbury CT2 7NR, Kent, England
[6] Lelintiz Inst Zoo & Wildlife Res IZW, Alfred Kowalke Str 17, D-10315 Berlin, Germany
[7] Nat Biodivers Ctr, Darwinweg 2,POB 9517, NL-2300 RA Leiden, Netherlands
[8] Univ Queensland, Sch Earth & Environm Sci, Ctr Biodivers & Conservat Sci, Brisbane, Qld 4072, Australia