[1] Univ Javeriana, Dept Ecol & Territorio, Fac Estudios Ambientales & Rurales, Bogota, DC, Colombia
[2] Ctr Ecol, Brisbane, Qld 4072, Australia
[3] Univ Queensland, Sch Geog Planning & Architecture, Ctr Remote Sensing & Informat Sci, St Lucia, Qld 4067, Australia
[4] Univ Queensland, Sch Geog Planning & Architecture, Ctr Ecol, St Lucia, Qld 4067, Australia