[1] INRAE, Inst Agro, IFREMER, DECOD Ecosyst Dynam & Sustainabil, F-44980 Nantes, France
[2] INRAE, Inst Agro, IFREMER, DECOD Ecosyst Dynam & Sustainabil, F-35042 Rennes, France
[3] Rennes Univ, Math Res Inst Rennes IRMAR, F-35042 Rennes, France
[4] INRAE, Inst Agro, IFREMER, DECOD Ecosyst Dynam & Sustainabil, F-29280 Brest, France
[5] NOAA, Habitat & Ecol Proc Res Program, Alaska Fisheries Sci Ctr, Natl Marine Fisheries Serv, Seattle, WA 98112 USA