[1] UMR Aix Marseille Univ, CNRS, IRD, Coll France,CEREGE, F-13545 Aix En Provence 4, France
[2] Dept Earth Sci, Oxford OX1 3AN, England
[3] Univ Tokyo, Atmosphere & Ocean Res Inst, Kashiwa, Chiba 2778564, Japan
[4] Univ Tokyo, Dept Earth & Planetary Sci, Kashiwa, Chiba 2778564, Japan
[5] Natl Inst Polar Res, Tachikawa, Tokyo 1908518, Japan
[6] JAMSTEC, Inst Biogeosci, Yokosuka, Kanagawa 2370061, Japan