[1] Fudan Univ, Sch Life Sci, Inst Biodivers Sci, Minist Educ,Key Lab Biodivers Sci & Ecol Engn,Coa, Shanghai 200438, Peoples R China
[2] Zhejiang Univ, Coll Life Sci, Lab Systemat & Evolutionary Bot & Biodivers, Hangzhou, Peoples R China
[3] East China Normal Univ, Sch Ecol & Environm Sci, Zhejiang Tiantong Forest Ecosyst Natl Observat &, Shanghai 200241, Peoples R China
[4] Inst Ecochongming IEC, Shanghai 202162, Peoples R China
[5] Inner Mongolia Univ, Sch Ecol & Environm, Minist Educ, Key Lab Ecol & Resource Use Mongolian Plateau, Hohhot 010021, Peoples R China
[6] Inner Mongolia Univ, Sch Ecol & Environm, Inner Mongolia Key Lab Grassland Ecol, Hohhot 010021, Peoples R China
[7] Univ Brunei Darussalam, Fac Sci, BE-1410 Gadong, Brunei