[1] Univ Geneva, Dept Neurosci Fondamentales, Rue Michel Servet 1, CH-1211 Geneva, Switzerland
[2] Harvard Med Sch, Dept Neurobiol, 220 Longwood Ave, Boston, MA 02115 USA
[3] Univ Rochester, Dept Brain & Cognit Sci, Rochester, NY 14627 USA
[4] UCL, Gatsby Computat Neurosci Unit, London, England