[1] Univ Putra Malaysia, Fac Agr, Dept Aquaculture, Serdang 43400, Selangor, Malaysia
[2] Univ Putra Malaysia, Inst Biosci, Serdang 43400, Selangor, Malaysia
[3] Univ Putra Malaysia, Fac Agr & Food Sci, Dept Anim Sci & Fishery, Bintulu 97008, Sarawak, Malaysia