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Minist Guangdong Prov Jointly Breeding Base South, Guangdong Prov Publ Lab New Microbial Applicat Te, Guangdong Key Lab Microbial Culture Collect & App, Guangdong State Key Lab Appl Microbiol,Guangdong, Guangzhou 510070, Guangdong, Peoples R China Minist Guangdong Prov Jointly Breeding Base South, Guangdong Prov Publ Lab New Microbial Applicat Te, Guangdong Key Lab Microbial Culture Collect & App, Guangdong State Key Lab Appl Microbiol,Guangdong, Guangzhou 510070, Guangdong, Peoples R China