[1] Univ Western Cape, Dept Biodivers & Conservat Biol, Private Bag X 17, ZA-7535 Bellville, South Africa
[2] Univ Western Cape, Anim Prod Inst, Agr Res Council, Private Bag X 17, ZA-7535 Bellville, South Africa
[3] Univ Cape Town, Dept Biol Sci, Private Bag X 3, ZA-7701 Rondebosch, South Africa
[4] Univ Western Cape, DST NRF Ctr Excellence Food Secur, Private Bag X 17, ZA-7535 Bellville, South Africa
[5] Anim Prod Inst, Agr Res Council, Private Bag X 2, ZA-0062 Irene, South Africa