[1] Univ Toulouse, INRAE, UMR DYNAFOR, F-31326 Castanet Tolosan, France
[2] Univ Montpellier, Univ Paul Valery Montpellier 3, SENS, IRD,CIRAD, Site St Charles,Route Mende, F-34199 Montpellier 5, France
[3] ECODEVELOPPEMENT, INRAE, F-84914 Avignon, France
[4] Inst Agro, INRAE, ESA, UMR 0980,BAGAP, F-35042 Rennes, France
[5] INRAE, DSLP, F-17450 St Laurent de la Pree, France