[1] Aarhus Univ, Sect Soil Fauna Ecol & Ecotoxicol, Dept Biosci, Vejlsovej 25, DK-8600 Silkeborg, Denmark
[2] Aarhus Univ, Dept Biosci, Arctic Res Ctr, Ny Munkegade 114, DK-8000 Aarhus C, Denmark
[3] Univ Aveiro, Dept Biol, P-3810193 Aveiro, Portugal
[4] Univ Aveiro, CESAM Ctr Environm & Marine Studies, P-3810193 Aveiro, Portugal
[5] Dept Biosci, Sect Marine Divers & Expt Ecol, Frederiksborgvej 399,Bldg B1-21, DK-4000 Roskilde, Denmark