[1] Inst Marine Resources & Ecosyst Studies, IMARES, Part Wageningen UR, NL-1970 AB Ijmuiden, Netherlands
[2] Tech Univ Denmark, DTU Aqua, Natl Inst Aquat Resources, DK-2920 Charlottenlund, Denmark
[3] IFREMER, French Res Inst Explorat Sea, STH, F-29280 Plouzane, France