[1] Univ Melbourne, Peter Doherty Inst Infect & Immun, Dept Microbiol & Immunol, Parkville, Vic, Australia
[2] Univ Melbourne, Walter & Eliza Hall Inst Med Res, Parkville, Vic, Australia
[3] Univ Melbourne, Dept Med Biol, Parkville, Vic, Australia
[4] Univ Melbourne, Ctr Epidemiol & Biostat, Melbourne Sch Populat & Global Hlth, Melbourne, Vic, Australia
[5] Duke NUS Med Sch, Singapore, Singapore