[1] Mem Univ Newfoundland, Fisheries & Marine Inst, St John, NF A1C 5R3, Canada
[2] Fisheries & Oceans Canada, Inst Maurice Lamontagne, 850 Route Mer, Mont Joli, PQ G5H 3Z4, Canada
[3] Northeast Fisheries Sci Ctr, Natl Marine Fisheries Serv, 166 Water St, Woods Hole, MA 02543 USA
[4] Marine & Freshwater Res Inst, Fornubudum 5, IS-220 Hafnafjordur, Iceland