More fossil woods from the Palaeogene of northern Kyushu, Japan

This paper describes five species of dicotyledonous fossil wood from the lower Oligocene Tsuyazaki Formation in Tsuyazaki, Fukuoka Prefecture, northern Kyushu: Rhus palaeojavanica (Anacardiaceae), Alnus scalariforme (Betulaceae), Hamamelis prejaponica (Hamamelidaceae), Magnoliaceoxylon palaeogenica (Magnoliaceae) and Sonneratia kyushuensis (Sonneratiaceae). This brings the number of species described from the Tsuyazaki locality to 19. Among these 19 species modern equivalents of all species, except for Sonneratia, occur in temperate to subtropical forests. Sonneratia is found today in mangrove vegetation of tropical to subtropical regions. The presence of Sonneratia may suggest a warmer climate in Kyushu during the early Oligocene.