[1] Univ Paris 06, CNRS, UMR 7621, Lab Oceanog Biol Banyulus,Observ Oceanol Banyuls, F-66650 Banyuls Sur Mer, France
[2] Univ W Florida, Ctr Environm Diagnost & Bioremediat, Pensacola, FL 32561 USA
[3] Univ Mediterranee, CNRS, INSU, UMR 6117,Ctr Oceanol Mareseille,LMGEM, F-13288 Marseille, France
[4] Univ Paris 06, CNRS, UMR 7621, F-66650 Banyuls Sur Mer, France