[1] Yokohama City Univ, Grad Sch Med, Ophthalmol & Visual Sci, Yokohama, Kanagawa, Japan
[2] Keio Univ, Lab Retinal Cell Biol, Tokyo, Japan
[3] Hokkaido Univ, Grad Sch Med, Sapporo, Hokkaido, Japan
[4] Keio Univ, Lab Retinal Cell Biol, Ophthalmol, Tokyo, Japan
[5] Keio Univ, Ophthalmol, Tokyo, Japan