[1] Univ Victoria, Dept Biochem & Microbiol, Victoria, BC V8W 3P6, Canada
[2] Univ Alberta, Dept Chem, Edmonton, AB T6G 2G2, Canada
[3] Univ Victoria, Dept Biol, Victoria, BC V8W 3N5, Canada
[4] Univ Victoria, Ctr Biomed Res, Victoria, BC V8W 3N5, Canada