[1] Univ Huelva, Dept Agroforestry Sci, Huelva, Spain
[2] Univ Porto CIIMAR, Ctr Interdisciplinar Invest Marinha & Ambiental, Matosinhos, Portugal
[3] Univ Porto, Inst Ciencias Biomed Abel Salazar ICBAS, Porto, Portugal
[4] Inst Recerca & Tecnol Agroalimentaries IRTA, Aquaculture Program, Rapita, Spain