[1] Taishan Med Univ, Sch Life Sci, Tai An 271018, Shandong, Peoples R China
[2] Shandong Agr Univ, Coll Life Sci, Tai An 271018, Shandong, Peoples R China
[3] Shandong Agr Univ, Coll Forestry, Tai An 271018, Shandong, Peoples R China
[4] Univ Georgia, Coll Vet Med, Dept Vet Biosci & Diagnost Imaging, Athens, GA 30602 USA