[1] Univ Leuven, KU Leuven, Rega Inst, Dept Microbiol Immunol & Transplantat, Leuven, Belgium
[2] Univ Calif Los Angeles, David Geffen Sch Med, Dept Biomath, Los Angeles, CA 90095 USA
[3] Univ Calif Los Angeles, David Geffen Sch Med, Dept Human Genet, Los Angeles, CA 90095 USA
[4] Univ Calif Los Angeles, Fielding Sch Publ Hlth, Dept Biostat, Los Angeles, CA USA
[5] Univ Arizona, Dept Ecol & Evolutionary Biol, Tucson, AZ USA