[1] Woods Hole Oceanog Inst, Dept Marine Chem & Geochem, Woods Hole, MA 02543 USA
[2] Univ So Calif, Dept Biol Sci, Los Angeles, CA 90089 USA
[3] Univ So Calif, Dept Earth Sci, Los Angeles, CA 90089 USA
[4] Univ Calif Berkeley, Lawrence Berkeley Lab, Berkeley, CA 94720 USA
[5] Bigelow Lab Ocean Sci, W Boothbay Harbor, ME USA