[1] Sultan Moulay Slimane Univ, Polydisciplinary Fac, Unit Biotechnol & Sustainable Dev Nat Resources B, Beni Mellal, Morocco
[2] Fac Sci & Tech, Unit Biotechnol & Symbiosis Agrophysiol, Marrakech, Morocco
[3] Georgia Inst Technol, Sch Math, Atlanta, GA 30332 USA
[4] Natl Inst Agron Res INRA, Unit Plant Breeding, Marrakech, Morocco
[5] Mohammed VI Polytech Univ, Agrobiosci AgBs Program, Benguerir, Morocco