[1] Beijing Forestry Univ, Sch Soil & Water Conservat, Beijing 100083, Peoples R China
[2] Beijing Forestry Univ, Sch Soil & Water Conservat, CNERN, Jixian Natl Forest Ecosyst Observat & Res Stn, Beijing, Peoples R China
[3] Chinese Acad Sci, Inst Tibetan Plateau Res, Key Lab Alpine Ecol, Beijing, Peoples R China
[4] Chinese Acad Sci, Inst Tibetan Plateau Res, Beijing, Peoples R China