[1] Univ Penn, Sch Med, Ctr Neurodegenerat Dis Res, Inst Aging, Philadelphia, PA 19104 USA
[2] Univ Penn, Sch Med, Dept Pathol & Lab Med, Philadelphia, PA 19104 USA
[3] Univ Penn, Sch Med, Dept Neurol, Philadelphia, PA 19104 USA
[4] Univ Calif San Francisco, Dept Neurol, San Francisco, CA 94143 USA