[1] Univ Catolica Norte, Fac Ciencias Mar, Biol & Ecol Aplicada, Coquimbo, Chile
[2] Univ Catolica Norte, Fac Ciencias Mar, Ctr Estudios Avanzados Zonas Aridas, Coquimbo, Chile
[3] Univ Bernardo Higgins, Ctr Invest Recursos Nat & Sustentabilidad, Santiago, Chile
[4] Oregon State Univ, Coll Earth Ocean & Atmospher Sci, Corvallis, OR 97331 USA
[5] Univ Austral Chile, Ctr FONDAP Invest Dinam Ecosistemas Marino Atlas, Valdivia, Chile
[6] Oregon State Univ, Dept Bot & Plant Pathol, Corvallis, OR 97331 USA