[1] Univ Witwatersrand, Sch Anim Plant & Environm Sci, ZA-2050 Johannesburg, South Africa
[2] Rochester Inst Technol, Chester F Carlson Ctr Imaging Sci, Rochester, NY 14623 USA
[3] Carnegie Inst, Dept Global Ecol, Stanford, CA 94305 USA
[4] Meraka Inst, CSIR, Remote Sensing Res Unit, ZA-0001 Pretoria, South Africa