[1] Chinese Acad Sci, State Key Lab Palaeobiol & Stratig, Nanjing Inst Geol & Palaeontol, Nanjing 210008, Peoples R China
[2] Chinese Acad Sci, Ctr Excellence Life & Paleoenvironm, Nanjing 210008, Peoples R China
[3] Univ Bristol, Sch Earth Sci, Life Sci Bldg,Tyndall Ave, Bristol BS8 1TQ, Avon, England
[4] Oregon State Univ, Oregon State Arthropod Collect, Dept Integrat Biol, Corvallis, OR 91331 USA
[5] Chinese Acad Sci, Inst Zool, Key Lab Zool Systemat & Evolut, Beijing 100101, Peoples R China
[6] Univ Chinese Acad Sci, Beijing 100049, Peoples R China
[7] Hainan Yazhou Bay Seed Lab, Bldg 1,7 Yiju Rd, Yazhou Dist 512024, Sanya, Peoples R China