[1] Univ Coimbra, Dept Life Sci, Ctr Funct Ecol, P-3000456 Coimbra, Portugal
[2] Univ Lisbon, Sch Agr, InBIO Associate Lab, Ctr Appl Ecol Prof Baeta Neves, P-1349017 Lisbon, Portugal
[3] Polytech Inst Coimbra, Sch Agr, P-3045601 Coimbra, Portugal
[4] Univ Porto, InBIO Associate Lab, Res Ctr Biodivers & Genet Resources, REN Biodivers Chair, Campus Agr Vairao, P-4485601 Vairao, Portugal