[1] Chinese Acad Sci, Inst Oceanol, Qingdao 266071, Peoples R China
[2] Chinese Acad Sci, Grad Sch, Beijing 100039, Peoples R China
[3] Ocean Univ China, Coll Marine Life Sci, Key Lab Marine Genet & Breeding, Chinese Minist Educ, Qingdao 266003, Peoples R China