[1] Univ Utrecht, Sect Ecol & Biodivers, Inst Environm Biol, Fac Sci, NL-3508 TB Utrecht, Netherlands
[2] UAB, Programa Manejo Bosques Amazonia Boliviana PROMAB, Riberalta, Bolivia
[3] Univ Sao Paulo, Inst Biociencias, BR-05508090 Sao Paulo, Brazil
[4] Univ Utrecht, Dept Phys & Astron, Fac Sci, NL-3508 TB Utrecht, Netherlands