[1] Univ Alabama, Dept Biol Sci, Tuscaloosa, AL 35487 USA
[2] Univ Georgia, Odum Sch Ecol, Athens, GA 30602 USA
[3] Univ Georgia, Savannah River Ecol Lab, Aiken, SC 29808 USA
[4] Cornell Univ, Dept Ecol & Evolutionary Biol, Ithaca, NY 14853 USA
[5] Miami Univ, Dept Biol, Oxford, OH 45056 USA
[6] Miami Univ, Grad Program Ecol Evolut & Environm Biol, Oxford, OH 45056 USA