Two conflicting stratigraphic schemes describe the Siberian Karginskii interstade (Oxygen Isotope Stage 3 equivalent) as having: 1) relatively stable climate with environments more similar to the full glaciation: or 2) variable climate with landscapes that more closely approximate contemporary ones. New data from continuous lake cores and a nearly continuous section from western Beringia (WB) suggest that both schemes are valid. Herb-dominated communities, possibly with isolated populations of Larix, characterized northern WB with only a slight shift from relatively warm to cool summers during the mid-interstade. In contrast, herb and shrub tundra, steppe, forest-tundra, and modern Larix forest occurred at various times in areas of southern WB, suggesting greater climatic instability. A thermal optimum is evident in the south during the mid-interstade, with modern vegetation in southeastern WB and Larix forest-tundra in the southwest. Variations in Pinus pumila pollen indicate summer warm/winter dry and summer warm/winter wet conditions in southeastern WB. These fluctuations contrast to other areas of WB, where summers and probably winters were consistently arid. Although the interstade presents a unique interval within the Late Pleistocene, paleodata and paleoclimatic models suggest that changes in marine conditions, including sea level, were likely key drivers in the regional climate history. (C) 2011 Elsevier Ltd. All rights reserved.