[1] Univ Sheffield, Sch Med, Acad Neurol Unit, Sheffield S10 2RX, S Yorkshire, England
[2] Univ Newcastle Upon Tyne, Inst Human Genet, Newcastle Upon Tyne NE1 3BZ, Tyne & Wear, England
[3] Univ Sheffield, Sch Med, Acad Unit Pathol, Sheffield S10 2RX, S Yorkshire, England