[1] Shizuoka Univ, Fac Agr, Dept Appl Biol Chem, Shizuoka 4228529, Japan
[2] Kagoshima Univ, Fac Agr, Lab Anim Reprod, Kagoshima 8900065, Japan
[3] Hyogo Med Univ, Dept Genet, Nishinomiya, Hyogo 6638131, Japan
[4] Osaka City Univ, Sch Med, Lab Anim Ctr, Abeno Ku, Osaka 5450051, Japan