New species of karst-dwelling Pselaphinae from southwestern China (Coleoptera: Staphylinidae)

Three cavernicolous species of the ant-loving beetle subfamily Pselaphinae are described based on the material collected in the dark zone of limestone caves in southwestern China. Batraxis liyunchuni sp. nov. and Nipponobythus alienoceps sp. nov. from Guizhou are the first and third cave-dwelling species of their respective genus, but they do not exhibit particular troglomorphy compared to epigean congeners. Zopherobatrus lusciosus sp. nov. from Chongqing, showing typical morphological adaptation to hypogeal lifestyle, is the second known representative of this obligate cavernicolous genus. In light of the discovery of a second species, the generic diagnosis of Zopherobatrus is modified accordingly.