[1] Univ Reunion, Inst Rech Dev IRD, UMR 9220 ENTROPIE,CNRS, Inst Rech Dev,IFREMER,Univ Nouvelle Caledonie, Noumea, New Caledonia
[2] Univ Reunion, Inst Francais Rech Exploitat Mer, UMR 9220 ENTROPIE,CNRS, Inst Rech Dev,IFREMER,Univ Nouvelle Caledonie, Noumea, New Caledonia
[3] Govt French Polynesia, Marine Resources Div, Papeete, French Polynesi, France