[1] Alfred Wegener Inst Polar & Marine Res, DE-27515 Bremerhaven, Germany
[2] ETH, Inst Geol, CH-8092 Zurich, Switzerland
[3] Univ Bremen, MARUM Ctr Marine Environm Sci, D-28359 Bremen, Germany
[4] Univ Concepcion, Fac Ciencias Nat & Oceanog, Dept Zool, Concepcion, Chile
[5] Inst Catalana Recerca & Estud Avancats, Barcelona, Spain
[6] Univ Autonoma Barcelona, Inst Ciencia & Tecnol Ambientals, Bellaterra, Spain