[1] Taizhou Univ, Inst Wetland Ecol & Clone Ecol, Zhejiang Prov Key Lab Plant Evolutionary Ecol & C, Taizhou 318000, Peoples R China
[2] Binzhou Univ, Shandong Key Lab Ecoenvironm Sci Yellow River Del, Binzhou 256603, Peoples R China
[3] Beijing Forestry Univ, Sch Ecol & Nat Conservat, Beijing 100083, Peoples R China
[4] Univ Massachusetts, Biol Dept, 611 North Pleasant St, Amherst, MA 01003 USA