[1] Newcastle Univ, Sch Biomed Sci, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England
[2] Newcastle Univ, Inst Cellular Med, Newcastle Upon Tyne NE1 7RU, Tyne & Wear, England
[3] Northumbria Univ, Dept Sport Exercise & Rehabil, Newcastle Upon Tyne, Tyne & Wear, England
[4] Northwest Univ, Sch Environm Sci & Dev, Water Res Grp, ZA-2520 Potchefstroom, South Africa