[1] Hokkaido Univ, Grad Sch Environm Sci, N10W5, Sapporo, Hokkaido 0600810, Japan
[2] Hokkaido Univ, Arctic Res Ctr, N21W11, Sapporo, Hokkaido 0010021, Japan
[3] Nagoya Univ, Grad Sch Bioagr Sci, Chikusa Ku, Nagoya, Aichi 4648601, Japan
[4] Japan Agcy Marine Earth Sci & Technol, Res & Dev Ctr Global Change, Kanazawa Ku, 3173-25 Showa Machi, Yokohama, Kanagawa 2360001, Japan
[5] Japan Agcy Marine Earth Sci & Technol, Inst Arctic Climate & Environm Res, Kanazawa Ku, 3173-25 Showa Machi, Yokohama, Kanagawa 2360001, Japan
[6] Hokkaido Univ, Fac Environm Earth Sci, N10W5, Sapporo, Hokkaido 0600810, Japan
[7] Russian Acad Sci, Siberian Branch, Inst Biol Problems Cryolithozone, 41 Lenin Ave, Yakutsk 677980, Russia
[8] North Eastern Fed Univ, Yakutsk 677000, Russia
[9] Hokkaido Univ, Global Inst Collaborat Res & Educ, Global Stn Arctic Res, N21W11, Sapporo, Hokkaido 0010021, Japan
[10] Nagoya Univ, Grad Sch Environm Studies, Chikusa Ku, Nagoya, Aichi 4648601, Japan