[1] Marine & Coastal Management, Dept Environm Affairs, ZA-8012 Cape Town, South Africa
[2] Univ Cape Town, Dept Zool, Anim Demog Unit, ZA-7701 Rondebosch, South Africa
[3] Univ Cape Town, Percy Fitzpatrick Inst, DST NRF Ctr Excellence, ZA-7701 Rondebosch, South Africa