[1] Lund Univ, Dept Phys Geog & Ecosyst Sci, Solvegatan 12, SE-22362 Lund, Sweden
[2] Tech Univ Denmark, Dept Environm Engn, Bygningstorvet 115, DK-2800 Lyngby, Denmark
[3] Swedish Univ Agr Sci, Dept Forest Ecol & Management, SE-90183 Umea, Sweden
[4] Malmo Univ, Dept Mat Sci & Appl Math, SE-20506 Malmo, Sweden