[1] Univ Calif San Diego, Scripps Inst Oceanog, La Jolla, CA 92093 USA
[2] Monterey Bay Aquarium Res Inst, Moss Landing, CA USA
[3] Inst Maurice Lamontagne, Mont Joli, PQ G5H 3Z4, Canada
[4] Univ Washington, Sch Oceanog, Seattle, WA 98195 USA
[5] CSIRO Marine & Atmospher Res, Hobart, Tas, Australia