[1] Scottish Assoc Marine Sci, Scottish Marine Inst, Oban PA37 1QA, Argyll, Scotland
[2] Plymouth Marine Lab, Plymouth PL1 3DH, Devon, England
[3] Univ Southampton, Ocean & Earth Sci, Natl Oceanog Ctr Southampton, Southampton SO14 3ZH, Hants, England
[4] British Geol Survey, Edinburgh EH9 3LA, Midlothian, Scotland
[5] Univ Southampton, Natl Oceanog Ctr, Southampton SO14 3ZH, Hants, England
[6] Univ Edinburgh, Grant Inst, Sch Geosci, Edinburgh EH9 3JW, Midlothian, Scotland
[7] Scottish Assoc Marine Sci, NERC Natl Facil Sci Diving, Oban PA37 1QA, Argyll, Scotland
[8] Zayed Univ, Dubai, U Arab Emirates