[1] Univ E Anglia, Tyndall Ctr Climate Change Res, Norwich NR4 7TJ, Norfolk, England
[2] Univ E Anglia, Sch Environm Sci, Norwich NR4 7TJ, Norfolk, England
[3] ETH, Inst Biogeochem & Pollutant Dynam, Environm Phys, CH-8092 Zurich, Switzerland
[4] Univ Manchester, Sch Earth Atmospher & Environm Sci, Manchester M13 9PL, Lancs, England
[5] NOAA, Pacific Marine Environm Lab, Seattle, WA 98115 USA
[6] Woods Hole Oceanog Inst, Dept Marine Chem & Geochem, Woods Hole, MA 02543 USA
[7] Univ Sud Toulon Var, Aix Marseille Univ, CNRS, INSU,IRD,MIO,UM110, F-13288 Marseille 09, France
[8] Natl Marine Fisheries Serv, Silver Spring, MD USA
[9] Univ Angers, LUNAM Univ, Lab Bioindicateurs Actuels & Fossiles, CNRS,LPGN BIAF,UMR 6112, F-49045 Angers 01, France