[1] Natl Inst Polar Res, Tokyo 1900014, Japan
[2] Hokkaido Univ, Grad Sch Environm Sci, Sapporo, Hokkaido 0600810, Japan
[3] Naruto Univ Educ, Coll Educ, Naruto 7728502, Japan
[4] Hokkaido Univ, Fac Environm Earth Sci, Sapporo, Hokkaido 0600810, Japan
[5] Nagoya Univ, Grad Sch Bioagr Sci, Nagoya, Aichi 4648601, Japan
[6] Russian Acad Sci, Inst Biol Problem Cryolithozone, Siberian Div, Yakutsk 678891, Russia
[7] North Eastern Fed Univ, Yakutsk 677000, Russia