[1] Diretoria Pesquisa Cient, Inst Pesquisas Jardim Bot Rio de Janeiro, BR-22460030 Rio De Janeiro, RJ, Brazil
[2] Univ Western Cape, Dept Biodivers & Conservat Biol, ZA-7535 Bellville, South Africa
[3] Smithsonian Inst, Natl Museum Nat Hist, Dept Bot, Washington, DC 20560 USA
[4] Univ Fed Rio de Janeiro, Ctr Ciencias Saude, Inst Biol, BR-21941900 Rio De Janeiro, RJ, Brazil