[1] Nagoya Univ, Grad Sch Sci, Div Mat Sci Phys, Chikusa Ku, Nagoya, Aichi 4648602, Japan
[2] Univ Tokyo, Grad Sch Arts & Sci, Dept Life Sci Biol, Meguro Ku, Tokyo 1538902, Japan
[3] Osaka Prefecture Univ, Grad Sch Sci, Dept Biol Sci, Osaka 5998531, Japan