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S China Agr Univ, Coll Nat Resources & Environm, Lab Plant Nematol, Dept Plant Pathol,Res Ctr Nematodes Plant Quarant, Guangzhou 510642, Guangdong, Peoples R China S China Agr Univ, Coll Nat Resources & Environm, Lab Plant Nematol, Dept Plant Pathol,Res Ctr Nematodes Plant Quarant, Guangzhou 510642, Guangdong, Peoples R China